The Moss Flora (Musci) of the Sündiken Mountains

Turk J Bot
30 (2006) 137-148
© TÜB‹TAK
Research Article
The Moss Flora (Musci) of the Sündiken Mountains
Filiz SAVARO⁄LU*, Süleyman TOKUR
Osmangazi University, Faculty of Science and Arts, Department of Biology, 26480 Eskiflehir - TURKEY
*E-mail: fbirgi@ogu.edu.tr
Received: 11.08.2005
Accepted: 22.11.2005
Abstract: In this study carried out in the Sündiken Mountains, 83 taxa were identified by the authors. These belong to 21 families
and 42 genera of bryophytes. Of these, 64 taxa are new for the B7 grid-square adopted by Henderson.
Key Words: Bryophytes, moss, flora, Sündiken Mountains
Sündiken Da¤lar›n›n Karayosunu (Musci) Floras›
Özet: Bu çal›flmada, araflt›rma alan› olan Sündiken Da¤lar›ndan toplanan karayosunu örnekleri de¤erlendirilerek 21 familya ya ait 42
cins ve bunlara ba¤l› 83 takson tan›mlanm›flt›r. Bunlardan 64 tanesi Henderson taraf›ndan benimsenen kareleme sistemine göre B7
karesi için yenidir.
Anahtar Sözcükler: Bryophyta, karayosunu, flora, Sündiken Da¤lar›.
Introduction
The study area, the Sündiken Mountains, is located in
the B7 grid square according to the system adopted by
Henderson & Prentice (1969) (Figure 1). No detailed
study into the moss flora in this area, that is, our study
area, has been conducted to date. The Sündiken
Mountains are situated between the east latitudes of
29º58'-32° 04' and the north latitudes of 39º 06'-40°
09' in relation to the north-west of Central Anatolia
(Figure 2). The Sündiken Mountains possess a transitional
feature in terms of their locations and climatic conditions
in relation to their location between the west of the
Marmara region and Northern Anatolia. The vegetation of
this region is represented by 4 different common types of
vegetation, namely Pinus brutia Ten., P. nigra Arn. subsp.
pallasiana (Lamb.) Holmboe, P. sylvestris L. and Quercus
cerris L. var. cerris (Ekim & Akman, 1990).
Due to the topographical features of the region, there
exist some differences between the east-west and the
north-south parts with respect to climatic conditions. On
the one hand, the region is influenced by the oceanic
climate of Northern Anatolia. On the other hand, it is
influenced by the semi-arid and extremely cold climate of
the Mediterranean. These, accordingly, have an impact
upon the vegetation occurring in this region. The region
has an Eastern Mediterranean precipitation range (Ekim
& Akman, 1990). By and large, the length of the
Sündiken Mountains located within the boundaries of the
city of Eskiflehir is approximately 50 and 60 km, while its
width varies between 20 and 25 km. The mountain range
stretches in an east-west direction.
It is hoped that this study will contribute to the
knowledge of moss flora in Turkey and could be useful as
a guide for future research.
Materials and Methods
The materials for this study were collected during
periodic field trips between 2000 and 2002 in different
seasons.
The identification of the samples was carried out
using various books on moss flora (Smith, 1980; Nyholm,
1981; Watson, 1981; Greven, 1995). In addition, the
status of the species for the study area and for Turkey
was determined by reviewing the related literature
(Çetin, 1988a; Uyar & Çetin, 2004; Kürschner & Erda¤,
137
The Moss Flora (Musci) of the Sündiken Mountains
1
2
3
4
5
6
7
8
9
42°
A
40°
B
38°
C
36°
26°
28°
30°
32°
34°
36°
38°
40°
42°
44°
Figure 1. The grid system which is adopted by Henderson & Prentice for Turkey (1:10,000,000)
Gökçekaya dam
Saka
iver
May›slar 1048
Kü
rya r
Sar›yar dam
Alap›nar
Karacaören
plü
Sakar›l›ca
eam
str
1678
S
Yar›mca
Bozda¤
Hekimda¤
U
Da¤küplü
N
Tafltepe
D
I
K
1685
E
Taycilar
Baflören
N
Tand›r
Türkmen hill
M
Sö¤ütçük
Özdenk
O
Yal›mkaya stream
Yal›mkaya
U
N
Uzunyatak hill
T
1786
A
I
A¤açhisar
N
den
Öz
Yk. Daniflment
Sündiken hill
1769
Ar›kaya
Dereköy
Mihal›çç›k
tta
am
Mu
1317
tre
Muttalip
lip
str
ESK‹fiEH‹R
S
ks
Afl. Daniflment
K›z›ldoruk hill
1052
Çukurören
Delikliçam hill
Mihal›çç›k stream
eam
0
4
8
12
16
20 km
Figure 2. Map of the study area (1:350,000)
2005). The list of species is arranged according to the
system described by Corley et al. (1981). In addition,
collection dates, collector number, altitude and habitat
are mentioned. Taxa new for the B7 grid square were
determined by reviewing the related literature
(Henderson & Prentice, 1969; Yücel & Tokur, 1989;
Yücel & Magil, 1997). New records for the B7 grid
square are marked with an asterisk.
The specimens were deposited within the herbarium
of the University of Osmangazi, Faculty of Science and
Arts, Department of Biology.
138
Floristic List
BRYOPSIDA
Polytrichaceae
Atrichum P.Beauv.
*1. A. undulatum (Hedw.) P.Beauv.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on soil, 26 August 2000, Savaro¤lu 583.
F. SAVARO⁄LU, S. TOKUR
Distribution: Turkey (A2, A3, A4, B6), Europe, Asia,
Algeria, N. America.
Polytrichum Hedw.
2. P. juniperinum Hedw.
B7 Eskiflehir: Sündiken Mountains, near Çatac›k,
Pinus nigra subsp. pallasiana forest, alt. 1650 m, on soil,
29 April 2001, Savaro¤lu 701, 576.
Distribution: Sub-cosmopolitan and widespread in
temperate regions of both hemispheres.
*3. P. strictum Menzies ex Brid.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on soil, 26 August 2000, Savaro¤lu 593.
Distribution: Turkey (A1, A2), Europe, Iceland,
Caucasus, Asia, N. America, Greenland, Antarctica.
Dicranaceae
Dicranum Hedw.
4. D. scoparium Hedw.
B7 Eskiflehir: Sündiken Mountains, near Ar›kaya,
Pinus nigra subsp. pallasiana- Quercus cerris var. cerris
forest, alt. 1180 m, on soil, 23 July 2000, Savaro¤lu
469, 600, 656.
Distribution: Sub-cosmopolitan.
5. D. tauricum Sapjegin
B7 Eskiflehir: Sündiken Mountains, A¤açhisar, Pinus
nigra subsp. pallasiana forest, alt. 1580 m, on tree trunk,
26 August 2000, Savaro¤lu 554.
Distribution: Turkey (A2, A3, B6, B7), Europe, N.
America, Greenland.
Dicranoweisia Lindb. ex Milde
*6. D. cirrata (Hedw.) Lindb. ex Milde
B7 Eskiflehir: Sündiken Mountains, near Çatac›k,
Pinus nigra subsp. pallasiana forest, alt. 1560 m, on soil,
29 April 2001, Savaro¤lu 698, 661.
Distribution: Turkey (A1, A2, A3, B6, C11, C12),
Europe, Caucasus, Himalayas, Mongolia, Algeria, Canary
Islands, Hawaii, N. America, Tasmania.
Dicranella (Mull.Hal.) Schimp.
*7. D. varia (Hedw.) Schimp.
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1460 m, on rock,
28 March 2001, Savaro¤lu 650.
Distribution: Turkey (A1, A2, A3, A4, B6, C11, C12),
Europe, Kashmir, China, Siberia, N. Africa, N. America,
Cuba, Jamaica, Haiti.
Ditrichaceae
Ditrichum Hampe.
*8. D. heteromallum (Hedw.) E.Britton
B7 Eskiflehir: Sündiken Mountains, near Karakütük,
Pinus nigra subsp. pallasiana forest, alt. 1538 m, on tree
trunk, 15 October 2000, Savaro¤lu 601, 594.
Distribution: Turkey (A1, A2, A3, A4,), Europe,
Iceland, N. America.
*9. D. subulatum Hampe
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1440 m, on soil,
28 March 2001, Savaro¤lu 655.
Distribution: Turkey (B6, B8), Europe, Mediterranean
region, Madeira, Tenerife.
Encalyptaceae
Encalypta Hedw.
*10. E. rhaptocarpa Schwägr.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, on tree trunk, 10 May 2000, Savaro¤lu 22.
Distribution: Turkey (A1, A2, C11), Europe,
Caucasus, Asia, N. America, N. Africa, Greenland, Hawaii.
11. E. streptocarpa Hedw.
B7 Eskiflehir: Sündiken Mountains, Sakar››l›ca, Pinus
nigra subsp. pallasiana forest, alt. 620 m, on rock, 10
May 2000, Savaro¤lu 148.
Distribution: Turkey (A1, A2, A3, B6, B7, C12),
Europe, Caucasus, N. America.
*12. E. vulgaris Hedw.
B7 Eskiflehir: Sündiken Mountains, Sakar››l›ca, Pinus
nigra subsp. pallasiana forest, alt. 560 m, on rock, 10
May 2000, Savaro¤lu 24, 147.
Distribution: Cosmopolitan.
Pottiaceae
Syntrichia Brid.
*13. S. mucronifolia (Schwägr.) Brid.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, on rock, 10 May 2000, Savaro¤lu 38b.
139
The Moss Flora (Musci) of the Sündiken Mountains
Distribution: Turkey (B6, C11), Europe, Caucasus, N.
Asia, China, N. America, Greenland.
14. S. princeps (De Not.) Mitt.
Distribution: Turkey (B6), Europe, Mediterranean.
Trichostomum Bruch
*21. T. branchydontium Bruch.
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1340 m, on rock,
28 March 2001, Savaro¤lu 642.
B7 Eskiflehir: Sündiken Mountains, Bozda¤, Quercus
cerris var. cerris-Q. pubescens forest, alt. 1240 m, on
soil, 10 May 2000, Savaro¤lu 51b.
Distribution: Turkey (A1, A2, B6, B7, C11, C12, C13,
C14), Europe, Asia, N. and S. America, New Zealand.
Distribution: Turkey (A1, A4, B7, C11, C12, C13),
Europe, Caucasus, Syria, China, Japan, N. and S. Africa,
New Zealand.
15. S. ruralis (Hedw). F.Weber & D.Mohr.
B7 Eskiflehir: Sündiken Mountains, Bozda¤, Quercus
cerris var. cerris forest, alt. 1240 m, on tree trunk, 10
May 2000, Savaro¤lu 78, 566.
Distribution: Cosmopolitan.
*16. S. subulata (Hedw.) F.Weber & D.Mohr var.
subulata
B7 Eskiflehir: Sündiken Mountains, near Çatac›k,
Pinus nigra subsp. pallasiana forest, alt. 1650 m, on soil,
29 April 2001, Savaro¤lu 597, 692.
Distribution: Cosmopolitan.
*17. S. subulata (Hedw.) F.Weber & D.Mohr var.
angustata (Schimp.) J.J.Amann
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1510 m, on soil,
28 March 2001, Savaro¤lu 455, 609, 658.
Distribution: Turkey (A1, A2, B6, B8, C11), Europe,
Asia, N. America, N. Africa.
*18. S. subulata (Hedw.) F.Weber & D.Mohr var.
graeffii Warnst.
B7 Eskiflehir: Sündiken Mountains, Bozda¤, Quercus
cerris var. cerris forest, alt. 1240 m, on soil, 10 May
2000, Savaro¤lu 71, 454, 629, 696.
Distribution: Turkey (A2, C11), W. and N. Britain.
Tortula Hedw.
*19. T. muralis Hedw.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, on rock, 10 May 2000, Savaro¤lu 38a.
Distribution: Cosmopolitan.
Tortella (Lindb.) Limpr.
*22. T. tortuosa (Hedw.) Limpr.
B7 Eskiflehir: Sündiken Mountains, Alap›nar, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1240 m, on rock, 17 June 2000, Savaro¤lu 406.
Distribution: Cosmopolitan.
Cinclidotus P.Beauv.
*23. C. fontinaloides (Hedw.) P.Beauv.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, near water, on rock, 26 August 2000,
Savaro¤lu 591.
Distribution: Turkey (C11), Europe, Asia, Caucasus,
Tibet, Algeria, Morocco, Tunisia.
Grimmiaceae
Schistidium Bruch & Schimp.
24. S. apocarpum (Hedw.) Bruch & Schimp.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on rock, 26 August 2000, Savaro¤lu 578.
Distribution: Cosmopolitan.
*25. S. confertum (Funck) Bruch & Schimp.
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1340 m, on rock,
28 March 2001, Savaro¤lu 644.
Distribution: Turkey (A1, A2, A4, B6, B8, B9, C13,
C14), Europe, Caucasus, N. and C. Asia, Japan, Ethiopia,
N. America.
Pottia Ehrh. ex Furnr.
Grimmia Hedw.
*20. P. commutata Limpr.
*26. G. decipiens (Schultz) Lindb.
B7 Eskiflehir: Sündiken Mountains, Bozda¤, Quercus
cerris var. cerris-Q. pubescens forest, alt. 1240 m, on
soil, 10 May 2000, Savaro¤lu 51a.
140
B7 Eskiflehir: Sündiken Mountains, Tand›r, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1400 m, on rock, 11 June 2000, Savaro¤lu 278.
F. SAVARO⁄LU, S. TOKUR
Distribution: Turkey (A1, A2, A4, B6, C11, C13),
Europe, Algeria, Tenerife, N. America.
*27. G. longirostris Hook.
Distribution: Turkey (B6), Europe, Asia, Japan, N.
Africa, Algeria, Egypt, Azores, N. America.
Bryum Hedw.
B7 Eskiflehir: Sündiken Mountains, Alap›nar, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1260 m, on rock, 17 June 2000, Savaro¤lu 385a.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, on wet soil, 10 May 2000, Savaro¤lu 11.
Distribution: Turkey (A1, A2), Europe, Asia, Africa, N.
America.
Distribution: Turkey (A1, A2), Europe, Asia, C. and S.
Africa, Canary Islands, America.
*28. G. ovalis (Hedw.) Lindb.
B7 Eskiflehir: Sündiken Mountains, Ar›kaya, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1200 m, on rock, 23 July 2000, Savaro¤lu 446,
385.
Distribution: Cosmopolitan.
29. G. pulvinata (Hedw.) Sm.
B7 Eskiflehir: Sündiken Mountains, Sö¤ütçük, alt.
1140 m, on rock, 17 June 2000, Savaro¤lu 428, 452.
Distribution: Cosmopolitan.
*30. G. trichophylla Grev.
B7 Eskiflehir: Sündiken Mountains, north sides,
Karakütük, Pinus sylvestris forest, alt. 1516 m, on rock,
15 October 2000, Savaro¤lu 358, 617.
Distribution: Cosmopolitan.
*31. G. dissimulata E.Maier
B7 Eskiflehir: Sündiken Mountains, Ar›kaya, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1200 m, on rock, 23 July 2000, Savaro¤lu 443.
Distribution: Turkey (A2), N. Britain, Portugal.
Funariaceae
Funaria Hedw.
32. F. hygrometrica Hedw.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, on oak trunk (=Quercus sp.), 10 May 2000,
Savaro¤lu 7.
Distribution: Cosmopolitan.
Bryaceae
Pohlia Hedw.
*33. P. melanodon (Brid.) J.Shaw
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1460 m, near
water, on soil, 28 March 2001, Savaro¤lu 646.
*34. B. alpinum Huds. ex With var. viride Husn.
*35. B. bornholmense Wink. & R.Ruthe
B7 Eskiflehir: Sündiken Mountains, Sakar››l›ca, Pinus
nigra subsp. pallasiana forest, alt. 420 m, on soil, 31 May
2000, Savaro¤lu 184.
Distribution: Turkey (A2), C. Europe, N. America.
*36. B. caespiticium Hedw.
B7 Eskiflehir: Sündiken Mountains, Ar›kaya, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1230 m, on rock, 23 July 2000, Savaro¤lu 479,
695.
Distribution: Cosmopolitan.
37. B. capillare Hedw.
B7 Eskiflehir: Sündiken Mountains, north sides,
Karakütük, Pinus sylvestris forest, alt. 1516 m, on rock,
15 October 2000, Savaro¤lu 610.
Distribution: Cosmopolitan.
*38. B. creberrimum Taylor
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on soil, 26 August 2000, Savaro¤lu 581.
Distribution: Turkey (A2, A4, B6, B8, B9, C11, C13),
Europe, N. Asia, N. America, Australia, New Zealand.
*39. B. imbricatum (Schwägr.) Bruch & Schimp.
B7 Eskiflehir: Sündiken Mountains, near Yukar›
Daniflment, alt. 1270 m, on wet soil, 11 June 2000,
Savaro¤lu 345.
Distribution: Turkey (A2, A4), Europe, Asia, N. Africa,
N. America, Australia, Antarctica.
*40. B. mildeanum Jur.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, near water, on soil, 10 May 2000, Savaro¤lu 9.
Distribution: Turkey (A1, A2, C11, C13, C14),
Europe, Asia, N. Africa.
141
The Moss Flora (Musci) of the Sündiken Mountains
*41. B. schleicheri DC.
B7 Eskiflehir: Sündiken Mountains, Tafll›burun forest
store, Pinus nigra subsp. pallasiana forest, alt. 1510 m,
near water, on soil, 10 June 2001, Savaro¤lu 709.
Distribution: Turkey (A4, B8, C13), Europe, Iran,
Syria, Asia, N. Africa.
*42. B. subelegans Kindb.
B7 Eskiflehir: Sündiken Mountains, Bozda¤, Quercus
cerris var. cerris forest, alt. 1240 m, on soil, 10 May
2000, Savaro¤lu 77.
Distribution: Turkey (A1, A2), Europe, N. America.
*43. B. intermedium (Brid.) Blandow
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, near water, on rock, 10 May 2000, Savaro¤lu
25.
Distribution: Turkey (C11, C12), Europe, Asia, N.
Africa, N. America, Greenland, Australia.
Plagiomnium T.Kop.
*47. P. undulatum (Hedw.) T.Kop.
B7 Eskiflehir: Sündiken Mountains, north sides,
Karakütük, Pinus sylvestris forest, alt. 1516 m, near
water, 15 October 2000, Savaro¤lu 618.
Distribution: Turkey (A1, A2, A3, A4, B6, C11, C13),
Europe, Asia, N.W. Africa.
Aulacomniaceae
Aulacomnium Schwägr.
*48. A. androgynum (Hedw.) Schwägr.
B7 Eskiflehir: Sündiken Mountains, Ar›kaya, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1240 m, on tree roots, 23 July 2000, Savaro¤lu
482.
Distribution: Turkey (A1, A2, B6, B8), Europe, Japan,
N. America.
Bartramiaceae
Mniaceae
Philonotis Brid.
Mnium Hedw.
49. P. calcarea (Bruch & Schimp.) Schimp.
*44. M. stellare Hedw.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, near water, on wet soil, 26 August 2000,
Savaro¤lu 577.
Distribution: Turkey (A1, A2, A4), Europe, Turkey,
Asia, N. America.
*45. M. thomsonii Schimp.
B7 Eskiflehir: Sündiken Mountains, north sides,
Karakütük, Pinus sylvestris forest, alt. 1516 m, near
water, on wet soil, 15 October 2000, Savaro¤lu 607.
Distribution: Turkey (B6), Europe, Faroes, Caucasus,
C. Asia, Siberia, China, N. America, Greenland.
Rhizomnium T.Kop.
*46. R. punctatum (Hedw.) T.Kop.
B7 Eskiflehir: Sündiken Mountains, north sides,
Karakütük, Pinus sylvestris forest, alt. 1516 m, near
water, 15 October 2000, Savaro¤lu 605.
Distribution: Turkey (A1, A2, A3, A4, B8), Europe,
Asia, N. America, Greenland.
142
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, forest road, Code. 309, Pinus sylvestris
forest, alt. 1430 m, on rock in stream, 11 August 2000,
Savaro¤lu 523.
Distribution: Turkey (A1, A2, A3, A4, B7, B10, C13),
Europe, Faroes, Asia, Algeria, N. America.
*50. P. rigida Brid.
B7 Eskiflehir: Sündiken Mountains, Ar›kaya, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1380 m, near water, 23 July 2000, Savaro¤lu 490.
Distribution: Turkey (A4, B5), Belgium, Switzerland,
Portugal, Italy, Sicily, Caucasus, Algeria.
*51. P. seriata Mitt.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, forest road, Code. 309, Pinus sylvestris
forest, alt. 1430 m, on rock in stream, 11 August 2000,
Savaro¤lu 542.
Distribution: Turkey (A1, A2, A4, B6), Europe,
Faroes, Caucasus, Iran, Siberia, N. Africa, Greenland.
F. SAVARO⁄LU, S. TOKUR
Orthotrichaceae
Distribution: Cosmopolitan.
Orthotrichum Hedw.
Leucodontaceae
*52. O. affine Schrad. ex Brid.
Leucodon Schwägr.
B7 Eskiflehir: Sündiken Mountains, Ar›kaya, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1200 m, on tree trunk, 23 July 2000, Savaro¤lu
448a, 457, 569.
Turkey (A1, A2, A3, A4, B6), Europe, Asia, N. Africa,
N. America.
53. O. lyellii Hook. & Taylor
B7 Eskiflehir: Sündiken Mountains, Ar›kaya, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1200 m, on rock, 23 July 2000, Savaro¤lu 444,
453.
Distribution: Turkey (A2, A4, B7, C11), Europe, Italy,
Caucasus, Algeria, Morocco, N. and S. America, Mexico.
54. O. rupestre Schleich. ex Schwägr.
B7 Eskiflehir: Sündiken Mountains, Alap›nar, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1260 m, on rock, 17 June 2000, Savaro¤lu 382,
451, 462, 564.
Distribution: Cosmopolitan.
*55. O. speciosum Nees
B7 Eskiflehir: Sündiken Mountains, Bozda¤, Quercus
cerris var. cerris-Q. pubescens forest, alt. 1240 m, on
tree trunk, 10 May 2000, Savaro¤lu 63, 354, 448b,
460.
Distribution: Turkey (A1, A2, C11), Europe, N. Africa,
N. and C. Asia, N. America, Greenland.
Ulota D.Mohr.
*56. U. hutchinsiae (Sm.) Hammar
B7 Eskiflehir: Sündiken Mountains, near Çatac›k,
Kömürce-Kapakl› road, Pinus nigra subsp. pallasiana
forest, alt. 1650 m, on rock, 29 April 2001, Savaro¤lu
691.
Distribution: Turkey (B6), Europe, Japan, N. America.
Fontinalaceae
Fontinalis Hedw.
*57. F. antipyretica Hedw.
B7 Eskiflehir: Sündiken Mountains, Çatac›k roadside,
Pinus nigra subsp. pallasiana forest, alt. 1340 m, on rock
in stream, 7 July 2001, Savaro¤lu 764.
*58. L. immersus Lindb.
B7 Eskiflehir: Sündiken Mountains, near Ar›kaya,
Pinus nigra subsp. pallasiana- Quercus cerris var. cerris
forest, alt. 1180 m, on rock, 23 July 2000, Savaro¤lu
462a.
Distribution: Turkey (A2, A3), Europe, Asia.
*59. L. sciuroides (Hedw.) Schwägr.
B7 Eskiflehir: Sündiken Mountains, Pinus nigra subsp.
pallasiana-Quercus cerris var. cerris–Juniperus excelsa
forest, alt. 1340 m, on rock, 26 August 2000, Savaro¤lu
431, 562b.
Distribution: Cosmopolitan.
Antitrichia Brid.
*60. A. curtipendula (Hedw.) Brid.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on soil, 26 August 2000, Savaro¤lu 585.
Distribution: Turkey (A2, A3, A4, B6, C11), Europe,
Caucasus, C. and S. Africa, N. America, Greenland.
Neckeraceae
Neckera Hedw.
*61. N. pumila Hedw.
B7 Eskiflehir: Sündiken Mountains, Pinus nigra subsp.
pallasiana-Quercus cerris var. cerris–Juniperus excelsa
forest, alt. 1340 m, on rock, 26 August 2000, Savaro¤lu
567.
Distribution: Turkey (A2), C. Europe, Madeira,
Canaries.
Thamniaceae
Thamnobryum Nieuwl.
*62. T. alopecurum (Hedw.) Nieuwl. ex Gangulee
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on rock, 26 August 2000, Savaro¤lu 592.
Distribution: Turkey (A2, A3, A4, C11), Europe,
Faroes, Asia, N. Africa, N. America.
143
The Moss Flora (Musci) of the Sündiken Mountains
Amblystegiaceae
Brachythecium Schimp.
Cratoneuron (Sull.) Spruce
*68. B. erythrorrhizon Schimp.
63. C. filicinum (Hedw.) Spruce var. atrovirens
(Brid.) Ochyra
B7 Eskiflehir: Sündiken Mountains, Pinus nigra subsp.
pallasiana-Quercus cerris var. cerris–Juniperus excelsa
forest, alt. 1340 m, on wet soil, 26 August 2000,
Savaro¤lu 571.
Distribution: Turkey (A1, A2, A4, B6, B7, B10, C11,
C12), Europe, Asia, N. America, N. Africa.
Palustriella Ochyra
64. P. commutata (Hedw.) Ochyra
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on rock in stream, 26 August 2000, Savaro¤lu
573.
Distribution: Turkey (A2, A3, B7, B8, B9, C13, C15),
Europe, C. Asia, N. America, N. Africa.
Leptodictyum (Schimp.) Warnst.
65. L. riparium (Hedw.) Warnst.
B7 Eskiflehir: Sündiken Mountains, Baflören, Pinus
nigra subsp. pallasiana forest, alt. 1070 m, on rock, 17
June 2000, Savaro¤lu 411
Distribution: Turkey (A2, B6, B7), Cosmopolitan.
Brachytheciaceae
Isothecium Brid.
*66. I. alopecuroides (Dubois) Isov.
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on rock, 26 August 2000, Savaro¤lu 589.
Distribution: Turkey (A1, A2, A3, A4, B6, C11, C13),
Europe, Caucasus, Canaries, Azores.
Homalothecium Schimp.
67. H. sericeum (Hedw.) Schimp.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, on rock, 10 May 2000, Savaro¤lu 29, 34,
562a.
Distribution: Cosmopolitan.
144
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1400 m, on tree
trunk, 28 March 2001, Savaro¤lu 660.
Distribution: Turkey (A1, A2, A4), Europe, Asia, N.
America.
*69. B. plumosum (Hedw.) Schimp.
B7 Eskiflehir: Sündiken Mountains, Yukar› Daniflment,
alt. 1370 m, on oak trunk (=Quercus sp.), 11 June
2000, Savaro¤lu 332.
Distribution: Cosmopolitan.
70. B. populeum (Hedw.) Schimp.
B7 Eskiflehir: Sündiken Mountains, Yukar› Daniflment,
alt. 1370 m, on rock, 11 June 2000, Savaro¤lu 335.
Distribution: Turkey (A1, A2, A4, B6, B7),
cosmopolitan
*71. B. velutinum (Hedw.) Schimp. var. velutinum
B7 Eskiflehir: Sündiken Mountains, Yar›mca, Quercus
cerris var. cerris forest, alt. 1280 m, on tree trunk, 11
June 2000, Savaro¤lu 357.
Distribution: Turkey (A1, A2, A3, B6, C12), Europe,
Asia, N. America.
*72. B. velutinum (Hedw.) Schimp. var. salicinum
(Schimp.) Mönk.
B7 Eskiflehir: Sündiken Mountains, Hekimda¤, alt.
1240 m, on oak trunk (=Quercus sp.), 10 May 2000,
Savaro¤lu 20, 478.
Distribution: Turkey (A1, A2, A3, B6, C12), Europe,
Asia, N. Africa.
*73. B. velutinum (Hedw.) Schimp. var. validum
C.Jens.
B7 Eskiflehir: Sündiken Mountains, near Yukar›
Daniflment, alt. 1270 m, on pine trunk (=Pinus sp.), 11
June 2000, Savaro¤lu 75, 347.
Distribution: Turkey (A2), Europe, Asia, Japan, N.
America.
Scleropodium Schimp.
*74. S. tourettii (Brid.) L.F.Koch
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on rock, 26 August 2000, Savaro¤lu 575.
F. SAVARO⁄LU, S. TOKUR
Distribution: Turkey (C11), Denmark, Belgium,
Luxemburg, France, Germany, N. America.
Platyhypnidium M.Fleisch.
*75. P. riparioides (Hedw.) Dixon
B7 Eskiflehir: Sündiken Mountains, north sides, near
Karakütük, monument forest, Pinus sylvestris forest, alt.
1420 m, on rock in stream, 26 August 2000, Savaro¤lu
582.
Distribution: Turkey (A1, A2, A3, A4, A5, B6, B8, B9,
C11, C12), Europe, Asia, N. Africa, N. and C. America.
Eurhynchium Schimp.
*80. H. cupressiforme Hedw. var. lacunosum Brid.
B7 Eskiflehir: Sündiken Mountains, Bozda¤-Sakar››l›ca
roadside, Pinus nigra subsp. pallasiana-Quercus cerris var.
cerris forest, alt. 560 m, on soil, 10 May 2000,
Savaro¤lu 139.
Distribution: Turkey (A1, A2, A3, A4, B6, C12),
Europe, N. America.
*81. H. cupressiforme Hedw. var. resupinatum
(Taylor) Schimp.
B7 Eskiflehir: Sündiken Mountains, Yukar› Daniflment,
alt. 1370 m, on soil, 11 June 2000, Savaro¤lu 339.
Distribution: Turkey (A1, A2, A4, C11), Europe.
*76. E. hians (Hedw.) Sande Lac.
B7 Eskiflehir: Sündiken Mountains, north sides,
Karakütük, Pinus sylvestris forest, alt. 1516 m, near
stream, on soil, 15 October 2000, Savaro¤lu 613.
Distribution: Turkey (A1, A2, A3, B6, C11, C13),
Europe, Caucasus, Asia, Japan, Algeria, N. America.
Entodontaceae
Entodon Mull.Hal.
*82. H. imponens Hedw.
B7 Eskiflehir: Sündiken Mountains, near Yal›mkaya,
Pinus nigra subsp. pallasiana forest, alt. 1400 m, on rock,
28 March 2001, Savaro¤lu 659.
Distribution: Turkey (A2, A3), Europe, C. Asia,
Switzerland, Iceland, Japan, N. America.
*83. H. jutlandicum Holmen & E.Warncke
*77. E. concinnus (De Not.) Paris
B7 Eskiflehir: Sündiken Mountains, north sides,
Karakütük, Pinus sylvestris forest, alt. 1516 m, near
water, on soil, 15 October 2000, Savaro¤lu 608.
B7 Eskiflehir: Sündiken Mountains, Sakar››l›ca
roadside, Pinus nigra subsp. pallasiana- Quercus cerris
var. cerris forest, alt. 560 m, on tree trunk, 10 May
2000, Savaro¤lu 110.
Distribution: Turkey (C11),
Caucasus, Iran, Asia, N. America.
Distribution: Turkey (A2), C. Europe, Algeria, N.
America.
Europe,
Iceland,
Plagiotheciaceae
Taxiphyllum M.Fleisch.
Results and Discussion
*78. T. wissgrillii (Garov.) Wijk & Margad.
In this study, 83 taxa belonging to 42 genera and 21
families were identified from 814 moss specimens
collected from the Sündiken Mountains between 2000
and 2002.
B7 Eskiflehir: Sündiken Mountains, Tand›r, Pinus
nigra subsp. pallasiana-Quercus cerris var. cerris forest,
alt. 1420 m, on tree root and base, 11 June 2000,
Savaro¤lu 304.
Distribution: Turkey (A2), Europe, Iceland, Caucasus.
Hypnaceae
Hypnum Hedw.
79. H. cupressiforme Hedw. var. cupressiforme
B7 Eskiflehir: Sündiken Mountains, A¤açhisar, Pinus
nigra subsp. pallasiana forest, alt. 1580 m, on tree trunk,
26 August 2000, Savaro¤lu 555.
Distribution: Turkey (A1, A2, A3, A4, A5, B6, B7,
C11, C13), cosmopolitan.
According to Henderson’s grid system, the number of
new records in the study area in the B7 square is 64.
Species such as Ditrichum subulatum, Pottia commutata,
Pohlia melanodon, Bryum intermedium, Mnium
thomsonii, Philonotis rigida and Entodon concinnus are
some of the rarest found in this country and these
localities are mentioned for the first time.
In Table 1, the families Pottiaceae, Bryaceae,
Brachytheciaceae, Grimmiaceae, Orthotrichaceae and
Hypnaceae make up 60.8% of the total taxa in the study
area, and the other 15 families constitute 39.2%.
145
The Moss Flora (Musci) of the Sündiken Mountains
Table 1. The distributions of the taxa according to the families.
Number
of taxa
Percentage of
taxa according to
the total number of taxa
Pottiaceae
11
13.1
Bryaceae
11
13.1
Brachytheciaceae
11
13.1
Grimmiaceae
8
9.52
Orthotrichaceae
5
5.95
Hypnaceae
5
5.95
Dicranaceae
4
4.76
Mniaceae
4
4.76
Polytrichaceae
3
3.57
Encalyptaceae
3
3.57
Bartramiaceae
3
3.57
Leucodontaceae
3
3.57
Amblystegiaceae
3
3.57
Ditrichaceae
2
2.38
Funariaceae
1
1.19
Aulacomniaceae
1
1.19
Fontinalaceae
1
1.19
Neckeraceae
1
1.19
Thamniaceae
1
1.19
Entodontaceae
1
1.19
Plagiotheciaceae
1
1.19
Bryopsida Families
Based on the observations made during the field trips,
acrocarpous mosses are capable of surviving during arid
periods despite exposure to the graining effects of
constant sunlight and the wind in arid habitats. On the
other hand, pleurocarpous mosses were observed to be
dominant in the habitats where water is abundant and the
air is relatively humid. In consideration of the whole study
area, 12 of the families belong to acrocarpous and 9
belong to pleurocarpous mosses. As to the taxa they
included, 67.46% (56 taxa) of the taxa occurring in the
Sündiken Mountains belong to acrocarpous mosses,
whilst 32.53% (27 taxa) belong to pleurocarpous
mosses. The fact that the region has a semi-arid
characteristic seems to account for such a distribution of
acrocarpous and pleurocarpous mosses.
Due to the altitude features of the study area, there
are some climatic differences between the directions of
east-west and north-south. On the one hand, the area is
exposed to the oceanic climate of Northern Anatolia. On
the other hand, it is exposed to the semi-arid and very
146
cold Mediterranean climate of Central Anatolia. This
feature seems to have an effect upon the vegetation
occurring in these areas. The best vegetation occurs in
Çatac›k-Karakütük environs, where the forest is the
densest. Therefore, mosses densely occur in this specific
area. Because the western part of the study area is less
rugged than its eastern part, the parts cleared of plants
or trees have been harnessed as agricultural areas.
Therefore, there is a spare distribution of mosses here.
In Table 2, a comparison between studies carried out
in the Black Sea (Çetin & Yurdakulol, 1985, 1988; Çetin
& Uyar, 1997) and the Mediterranean (Çetin, 1989)
shows that the proportion of acrocarpous mosses in the
Mediterranean is higher than that in the Black Sea region.
In particular, the number of taxa of Pottiaceae family
members is higher than the number of taxa of
Brachytheciaceae family members in the study areas, due
to the reduced forest in the study areas because of
excessive construction. Consequently, pleurocarpous taxa
decreased since these taxa need much more humidity than
acrocarpous taxa such as Pottiaceae family members,
which can endure long periods of heat and dryness.
In the north parts of the study area (1300-1700 m)
covered by Pinus sylvestris forest, there occur
Polytrichum strictum, P. juniperinum, Atrichum
undulatum, Bryum creberrimum, Antitrichia curtipendula,
Dicranoweisia cirrata, Dicranum scoparium, Syntrichia
subulata var. subulata, S. subulata var. angustata, S.
subulata var. graeffii, Mnium stellare, M. thomsonii and
Rhizomnium punctatum on the acidic soil; Schistidium
apocarpum, Grimmia trichophylla, Bryum capillare, B.
caespiticium, Thamnobryum alopecurum, Isothecium
myurum, Scleropodium tourettii, Ulota hutchinsiae and
Cinclidotus fontinaloides on rocks; Ditrichum
heteromallum on tree trunks; Bryum schleicheri var.
latifolium, Mnium thomsonii, Rhizomnium punctatum,
Plagiomnium undulatum, Philonotis seriata, P. calcarea,
Palustriella commutata, Rhynchostegium riparioides,
Eurhynchium hians and Entodon concinnus in wetlands
and near water in places; and Fontinalis antipyretica in
water.
Likewise, in the north parts of the study area (11001300 m) covered by Pinus nigra subsp. pallasianaQuercus cerris var. cerris forest, there occur Tortella
tortuosa, Grimmia affinis, G. ovalis and Orthotrichum
rupestre on rocks.
F. SAVARO⁄LU, S. TOKUR
Table 2. The comparision of the taxa distribution according to the families.
Sündiken
Mountains
Families
Bolu
Gerede-Aktafl
Yedigöller
National Park
Antalya and
Its Environs
Sinop and
Its Environs
The number
of taxa
%
The number
of taxa
%
The number
of taxa
%
The number
of taxa
%
The number
of taxa
%
Pottiaceae
11
13.1
16
12
12
13.6
32
35.5
12
10.7
Bryaceae
11
13.1
17
12.6
2
2.2
4
4.4
6
5.3
Brachytheciaceae
11
13.1
19
15
15
17.4
14
15.5
23
20.5
Grimmiaceae
8
9.5
8
6
3
3.4
6
6.6
3
2.6
Orthotrichaceae
5
6
5
3.8
2
2.2
8
8.9
5
4.5
Hypnaceae
5
6
11
8
8
9..9
3
3.3
10
8.9
In the west parts of northern regions of the area
(400-700 m) covered by Pinus nigra subsp. pallasiana
forest, there occur Bryum bornholmense and Hypnum
cupressiforme var. lacunosum on the soil, Encalypta
streptocarpa and E. vulgaris on rocks, and Hypnum
jutlandicum on tree trunks and roots.
In the south parts of the area (1000-1600 m)
covered by the Pinus nigra subsp. pallasiana-Quercus
cerris var. cerris-Juniperus excelsa forest, there occur
Dicranum
scoparium,
Ditrichum
subulatum,
Dicranoweisia cirrata, Syntrichia subulata var. angustata
and Pohlia melanodon on the soil; Grimmia pulvinata, G.
ovalis, G. trichophylla, G. dissimulata, Bryum
caespiticium, Amblystegium riparium, Orthotrichum
lyellii, O. rupestre, Leucodon immersus, L. sciuroides,
Homalothecium sericeum, Dicranella varia, Syntrichia
subulata var. graeffii, S. princeps, S. ruralis, Schistidium
confertum, Hypnum imponens and Neckera pumila on
rocks; Aulacomnium androgynum, Orthotrichum affine,
O. speciosum, Dicranum tauricum, Brachythecium
erythrorrhizon, B. velutinum var. salicinum and Hypnum
cupressiforme var. cupressiforme on tree trunks and
roots; and Philonotis rigida and Cratoneuron filicinum
var. atrovirens near water.
Leaf cells of Grimmia dissimulata are incrassate and all
cell walls are sinuose. They most often occur over acidic
rocks in the form of fuzzy dark coloured tufts. With this
feature of theirs, they differ G. trichophylla. They were
recorded in Turkey for the second time (Uyar & Çetin,
2000).
In the west part of the region (1200-1500 m)
covered by Quercus cerris var. cerris-Q. pubescens forest,
there occur Hypnum cupressiforme var. resupinatum,
Pottia commutata, Trichostomum branchydontium,
Bryum subelegans, B. alpinum var. viride and B.
mildeanum on soil; Grimmia decipiens, Brachythecium
populeum, Tortula muralis, Syntrichia mucronifolia,
Homalothecium sericeum and Encalypta vulgaris on
rocks; Taxiphyllum wissgrillii, Brachythecium velutinum
var. velutinum, B. velutinum var. validum, B. velutinum
var. salicinum, B, plumosum, Encalypta rhaptocarpa,
Syntrichia ruralis, S. subulata var. graeffii, Funaria
hygrometrica,
Orthotrichum
speciosum
and
Homalothecium sericeum on tree trunks and roots; and
Bryum inclinatum, B. alpinum var. viride and B.
intermedium in wetlands and near water in places.
Bryum alpinum var. viride have green leaves with
wide, long and thin walled cells. Border cells do not differ
from the other cells. Their leaf tips are denticulate. With
this feature of theirs, they differ from B. alpinum.
Brachythecium velutinum var. validum have larger
leaves and the surface of the seta is papillose with slightly
papillose spores. With this feature of theirs, they differ
from other varieties of B. velutinum. They were recorded
in Turkey for the second time (Uyar & Çetin, 2001b).
Acknowledgement
I would like to thank Prof. Dr. Barbaros Çetin for
identifying some of the species.
147
The Moss Flora (Musci) of the Sündiken Mountains
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